terricolous

Ferns were the first terricolous vascular plants, and still comprise an important part of primary productivity of ecosystems.

蕨类植物是最早登陆的维管植物，至今仍然是生态系统初级生产力的重要组成部分。

The vertical variation is that there are many autogenic carbonate sediments in the earlier period and that terricolous sediments are dominant in the later period.

纵向上呈现出由早期有较多内源碳酸盐沉积到晚期以大量陆源物为主的变化。